Tertiary Education:

1963 BSc University of Canterbury, New Zealand
1967 MSc (First Class Honours) University of Canterbury, New Zealand.
1971 PhD University of Canterbury, New Zealand


1968–1970 University Grants Committee Postgraduate Scholarship
1968–1970 N.Z. Internal Affairs Wildlife Scholarship
1976–1987 Appointed to Scientific Committee of Fiordland National Park Board
1980–1987 Appointed to Scientific Committee of New Zealand National Parks Authority
1983–1987 Appointed to Wapiti Scientific Advisory Committee by Minister of Lands
1989 J.S.Watson Conservation Award
1990 Co-convenor with C.M Perrins of symposium “Long-term population studies of birds” 20th International Ornithological Congress
1990 Prince and Princess of Wales Fellowship
1991 Senior Fulbright Fellowship- Cornell University


1971–1985 Scientist, N.Z. Wildlife Service, Wellington
1985–1987 Assistant Director of Research (Endangered Species), New Zealand Wildlife Service, Department of Internal Affairs,Wellington, N.Z
1987–1993 Scientist, Science & Research Directorate, Department of Conservation, Wellington, New Zealand
1991 Visiting Fellow, Section of Ecology & Systematics, Cornell University, Ithaca, New York, USA
1992 Visiting Fellow, Royal Ontario Museum, Toronto, Canada
1993–2012 Independent researcher
1996–1999 Marsden Grant, Royal Society of New Zealand